Nathan Fillion completed his school level education from the Holy Trinity Catholic High School, Concordia and graduation from the University College of Alberta and University, Alberta. Prior to shifting to the New York City in 1994, he actively participated in improv theatre which included Theatre sports with improvised soap opera Die-Nasty and Rapid Fire Theatre. He also starred in a TV Movie called Ordeal in the Arctic in 1993 alongside actors like Richard Chamberlain and also starred in Strange and Rich.
Fillion's eventually found his first recurring role in a daytime soap opera titled One Life to Live. His role as Joey Buchanan was well applauded his role also earned him the nomination for a Daytime Emmy Award in 1996. Fillion garnered huge critical acclaim and a massive fan following when he portrayed the role of Captain Malcolm Reynolds on the Joss Whedon's Firefly in 2002. He also made guest star appearances on various episodes of two short-lived TV shows, Miss Match, as Adam Logan and Pasadena, as Rev. Glenn Collins.
In 2009, he was cast to play the lead role as Richard Castle in ABC's hit television series, Castle. The show has lasted for more than 160 episodes and Nathan Fillion for his noteworthy performance won four People's Choice Awards for Favourite Dramatic TV Actor as of 2016. In addition to starring on "Castle", he has simultaneously appeared in several films like Super, Much Ado About Nothing, Percy Jackson: Sea of Monsters and in Thrilling Adventure Hour Live.